diff --git a/scene/resources/2d/skeleton/skeleton_modification_2d_lookat.cpp b/scene/resources/2d/skeleton/skeleton_modification_2d_lookat.cpp
index e5ce028dcc0..78335454a3f 100644
--- a/scene/resources/2d/skeleton/skeleton_modification_2d_lookat.cpp
+++ b/scene/resources/2d/skeleton/skeleton_modification_2d_lookat.cpp
@@ -141,37 +141,32 @@ void SkeletonModification2DLookAt::_execute(float p_delta) {
 		return;
 	}
 
-	Transform2D operation_transform = operation_bone->get_global_transform();
-	Transform2D target_trans = target_node_reference->get_global_transform();
+	real_t angle_to_target = operation_bone->get_angle_to(target_node_reference->get_global_position());
 
-	// Look at the target!
-	operation_transform = operation_transform.looking_at(target_trans.get_origin());
-	// Apply whatever scale it had prior to looking_at
-	operation_transform.set_scale(operation_bone->get_global_scale());
-
-	// Account for the direction the bone faces in:
-	operation_transform.set_rotation(operation_transform.get_rotation() - operation_bone->get_bone_angle());
+	// Account for the direction the bone faces in
+	angle_to_target -= operation_bone->get_bone_angle();
 
 	// Apply additional rotation
-	operation_transform.set_rotation(operation_transform.get_rotation() + additional_rotation);
+	angle_to_target += additional_rotation;
 
-	// Apply constraints in globalspace:
-	if (enable_constraint && !constraint_in_localspace) {
-		operation_transform.set_rotation(clamp_angle(operation_transform.get_rotation(), constraint_angle_min, constraint_angle_max, constraint_angle_invert));
-	}
+	if (enable_constraint) {
+		real_t new_angle = angle_to_target;
 
-	// Convert from a global transform to a local transform via the Bone2D node
-	operation_bone->set_global_transform(operation_transform);
-	operation_transform = operation_bone->get_transform();
-
-	// Apply constraints in localspace:
-	if (enable_constraint && constraint_in_localspace) {
-		operation_transform.set_rotation(clamp_angle(operation_transform.get_rotation(), constraint_angle_min, constraint_angle_max, constraint_angle_invert));
+		if (constraint_in_localspace) {
+			new_angle += operation_bone->get_rotation();
+			new_angle = clamp_angle(new_angle, constraint_angle_min, constraint_angle_max, constraint_angle_invert);
+			operation_bone->set_rotation(new_angle);
+		} else {
+			new_angle += operation_bone->get_global_rotation();
+			new_angle = clamp_angle(new_angle, constraint_angle_min, constraint_angle_max, constraint_angle_invert);
+			operation_bone->set_global_rotation(new_angle);
+		}
+	} else {
+		operation_bone->rotate(angle_to_target);
 	}
 
 	// Set the local pose override, and to make sure child bones are also updated, set the transform of the bone.
-	stack->skeleton->set_bone_local_pose_override(bone_idx, operation_transform, stack->strength, true);
-	operation_bone->set_transform(operation_transform);
+	stack->skeleton->set_bone_local_pose_override(bone_idx, operation_bone->get_transform(), stack->strength, true);
 }
 
 void SkeletonModification2DLookAt::_setup_modification(SkeletonModificationStack2D *p_stack) {
diff --git a/scene/resources/2d/skeleton/skeleton_modification_2d_twoboneik.cpp b/scene/resources/2d/skeleton/skeleton_modification_2d_twoboneik.cpp
index 8b4a6fcd095..2c60f620be8 100644
--- a/scene/resources/2d/skeleton/skeleton_modification_2d_twoboneik.cpp
+++ b/scene/resources/2d/skeleton/skeleton_modification_2d_twoboneik.cpp
@@ -142,7 +142,7 @@ void SkeletonModification2DTwoBoneIK::_execute(float p_delta) {
 		return;
 	}
 
-	// Adopted from the links below:
+	// Adapted from the links below:
 	// http://theorangeduck.com/page/simple-two-joint
 	// https://www.alanzucconi.com/2018/05/02/ik-2d-2/
 	// With modifications by TwistedTwigleg
@@ -150,9 +150,10 @@ void SkeletonModification2DTwoBoneIK::_execute(float p_delta) {
 	float joint_one_to_target = target_difference.length();
 	float angle_atan = target_difference.angle();
 
-	float bone_one_length = joint_one_bone->get_length() * MIN(joint_one_bone->get_global_scale().x, joint_one_bone->get_global_scale().y);
-	float bone_two_length = joint_two_bone->get_length() * MIN(joint_two_bone->get_global_scale().x, joint_two_bone->get_global_scale().y);
+	float bone_one_length = joint_one_bone->get_length() * MIN(joint_one_bone->get_global_scale().abs().x, joint_one_bone->get_global_scale().abs().y);
+	float bone_two_length = joint_two_bone->get_length() * MIN(joint_two_bone->get_global_scale().abs().x, joint_two_bone->get_global_scale().abs().y);
 	bool override_angles_due_to_out_of_range = false;
+	bool same_scale_sign = true;
 
 	if (joint_one_to_target < target_minimum_distance) {
 		joint_one_to_target = target_minimum_distance;
@@ -165,6 +166,10 @@ void SkeletonModification2DTwoBoneIK::_execute(float p_delta) {
 		override_angles_due_to_out_of_range = true;
 	}
 
+	if (joint_one_bone->get_global_scale().sign().x != joint_one_bone->get_global_scale().sign().y) {
+		same_scale_sign = false;
+	}
+
 	if (!override_angles_due_to_out_of_range) {
 		float angle_0 = Math::acos(((joint_one_to_target * joint_one_to_target) + (bone_one_length * bone_one_length) - (bone_two_length * bone_two_length)) / (2.0 * joint_one_to_target * bone_one_length));
 		float angle_1 = Math::acos(((bone_two_length * bone_two_length) + (bone_one_length * bone_one_length) - (joint_one_to_target * joint_one_to_target)) / (2.0 * bone_two_length * bone_one_length));
@@ -177,12 +182,23 @@ void SkeletonModification2DTwoBoneIK::_execute(float p_delta) {
 		if (std::isnan(angle_0) || std::isnan(angle_1)) {
 			// We cannot solve for this angle! Do nothing to avoid setting the rotation (and scale) to NaN.
 		} else {
-			joint_one_bone->set_global_rotation(angle_atan - angle_0 - joint_one_bone->get_bone_angle());
+			if (same_scale_sign) {
+				joint_one_bone->set_global_rotation(angle_atan - angle_0 - joint_one_bone->get_bone_angle());
+			} else {
+				joint_one_bone->set_global_rotation(angle_atan + angle_0 + joint_one_bone->get_bone_angle());
+			}
+
 			joint_two_bone->set_rotation(-Math::PI - angle_1 - joint_two_bone->get_bone_angle() + joint_one_bone->get_bone_angle());
 		}
+
 	} else {
-		joint_one_bone->set_global_rotation(angle_atan - joint_one_bone->get_bone_angle());
-		joint_two_bone->set_global_rotation(angle_atan - joint_two_bone->get_bone_angle());
+		if (same_scale_sign) {
+			joint_one_bone->set_global_rotation(angle_atan - joint_one_bone->get_bone_angle());
+			joint_two_bone->set_global_rotation(angle_atan - joint_two_bone->get_bone_angle());
+		} else {
+			joint_one_bone->set_global_rotation(angle_atan + joint_one_bone->get_bone_angle());
+			joint_two_bone->set_global_rotation(angle_atan + joint_two_bone->get_bone_angle());
+		}
 	}
 
 	stack->skeleton->set_bone_local_pose_override(joint_one_bone_idx, joint_one_bone->get_transform(), stack->strength, true);
@@ -211,7 +227,8 @@ void SkeletonModification2DTwoBoneIK::_draw_editor_gizmo() {
 	}
 	stack->skeleton->draw_set_transform(
 			stack->skeleton->to_local(operation_bone_one->get_global_position()),
-			operation_bone_one->get_global_rotation() - stack->skeleton->get_global_rotation());
+			operation_bone_one->get_global_rotation() - stack->skeleton->get_global_rotation(),
+			operation_bone_one->get_global_scale());
 
 	Color bone_ik_color = Color(1.0, 0.65, 0.0, 0.4);
 #ifdef TOOLS_ENABLED